Another pretty clear indication that it was used during the war is the white "blackout stripe" rather sloppily painted on the rear fender (mudguard, for the Brits), apparently in house paint. I can imagine the original owner cringing a bit as he slopped white paint on his glossy black fender and gold pinstripes.
The final clue, and the one that is perhaps the most moving for me, is that replacement brake pads appear to have been cut from salvaged rubber and installed in the shoes, I'm guessing in order to conserve resources during the war.
The sad part, and the real conundrum for folks like me, is that all of these features have now either been removed or covered up in the process of making the bike useable again. The material evidence of the bicycle's eventful past has literally been stripped away, out of necessity of course, but it's still a little sad. My great comfort, though, is that the bicycle will again be used for its intended purpose, rather than rusting in a back yard. I will also keep the tyres and brakes, of course, to pass on to the next owner of the Runwell, but hopefully that won't be for a long, long time.